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During the 2011 International Championship of Collegiate A Cappella (ICCA) Finals at the Lincoln Center, Barden University's all-female a cappella group, the Barden Bellas, loses to their all-male rival group, the Barden Treblemakers, due to a very nervous junior member Aubrey Posen projectile vomiting onstage in the middle of her solo under the pressure of her peers. Four months later, aspiring music producer Beca Mitchell enrolls as a freshman at Barden University at the insistence of her father. To her father's chagrin, Beca spends her time making mash-up mixes of songs and gets an internship at the school radio station, where she befriends fellow freshman Jesse Swanson.

At the university's activities fair, now-seniors Aubrey and Chloe Beale are attempting to recruit new members for the Bellas but find that interest in their group has decreased significantly after last year's on-stage incident. Beca is invited to join but declines and claims she can't sing. Later, Chloe hears Beca sing in the shower and convinces her to audition. At auditions, Beca sings a rendition of "Cups (When I'm Gone)", earning herself a place in the Bellas alongside fellow newcomers: tough Cynthia Rose Adams, promiscuous Stacie Conrad, eccentric Lilly Onakuramara, humorous Patricia "Fat Amy" Hobart, as well as Jessica Smith, Ashley Jones, and Denise. Meanwhile, Jesse joins the Treblemakers.